"Theory of Ancestral Contributions in Heredity," handwritten manuscript by Karl Pearson, published in Proceedings of the Royal Society (vol. 81:547) (5)

1975. [handwritten] 5 stantially modifies the offspring; there exists an "ancestral contribution" to the heritage. [tabular material] Grandparents & Offspring DD & DR & RR & + & - & DD & DR & RR 4 & -- & -- & 4 -- & 1 & -- & -- 3 & 1 & -- & 4 & -- & 6 & 2 & -- 3 & -- & 1 & 3 & 1 & 2 & 2 & -- 2 & 2 & -- & 4 & -- & 13 & 10 & 1 2 & 1& 1 & 3 & 1 & 8 & 14 & 2 2 & -- & 2 & 2 & 2 & 1 & 4 & 1 1 & 3 & -- & 4 & -- & 12 & 16 & 4 1 & 2 & 1 & 3 & 1 & 10 & 28 & 10 1 & 1 & 2 & 2 & 2 & 2 & 14 & 8 1 & -- & 3 & 1 & 3 & -- & 2 & 2 -- & 4 & -- & 4 & -- & 4 & 8 & 4 -- & 3 & 1 & 3 & 1 & 4 &16 &12 --& 2 & 2 & 2 & 2&1 & 10 & 13 -- & 1 & 3 & 1 & 3 -- & 2 & 6 -- & -- & 4 & -- & 4 & -- & -- & 1 [end tabular material] Of course the gametic contribution RR follows precisely the same system of parentages, and is again influenced by ancestry. If, however, we take the gametic contribution DR in such a population we find: